NBIK14010U Social Behaviour and Communication

Volume 2021/2022

MSc Programme in Biology
MSc Programme in Biology with a minor subject


The course covers the main topics in social behaviour and communication for animals, illustrated both by textbook examples and recent findings in the field; Levels of socialisation; Motivation, emotions and mood; Game theory and evolutionarily stable strategies; Communication networks and neighbourhood dynamics; Social cognition; Social aspects of animal welfare; Social and territorial behaviour; Signalling, reception and interpretation of communicative behaviour under constraints; Signals, information and meaning; Mating systems.

Learning Outcome


By completing the course the student will have obtained detailed knowledge of behavioural and communicative processes fundamental for animal social behaviour and organisation.


By completing the course the student will know how to:

  • Explain animal social behaviour and organisation in terms of function, mechanism, ontogeny and evolution;
  • Present in details the connecting and organising function of signals for animal social behaviour and explain the evolutionary forces that shape signals and signalling;
  • Explain the relative effects of physical, biological and social environments on individual behaviour.

By completing the course the student can:

  • Propose alternative hypotheses to explain a behaviour, design an appropriate experiment to test them and suggest predictions;
  • Measure behaviour using different methods and analyse the data obtained in different manners;
  • Critically discuss results with respect to the hypothesis tested, place them in a broad general context, link to other behaviour issues and propose further lines of enquiry;
  • Explain the costs and benefits of different experimental designs, data collection and analysis methods and identify sources of error;
  • Present her/his work (in oral and written form) at a level approaching the scientific standard;
  • Extract, present and critically discuss in detail the results of a scientific article about animal behaviour;
  • Identify and explain strengths and weaknesses in scientific articles, suggest solutions to the weaknesses and further enquiries inspired by the strengths.

Textbook and scientific articles (see Absalon for details).

We recommend that the course participants have basic knowledge about evolutionary biology (for example obtained in the course "Evolutionsbiologi").

Academic qualifications equivalent to a BSc degree is recommended.
The course includes a combination of plenary lectures given by the course leader and guest teachers who are experts in the field, discussions of recently-published scientific articles, and experiments using specialised software for measuring behaviour and analysing vocalisations as well as hand-on statistical analyses (notably in R software); two lectures and two discussion confrontations (colloquia) of two hours each plus one practical of four hours per week, including four group exercise reports and one oral presentation (also in group), for a total of six weeks, followed by an individual project consisting of a five page essay during the seventh week of the course. The course ends with an individual oral defence of the project and an oral examination on the rest of the course contents. All students must hand in at least three of the four exercise reports to take the exam.
  • Category
  • Hours
  • Class Instruction
  • 33
  • Preparation
  • 121,5
  • Practical exercises
  • 27
  • Project work
  • 20
  • Guidance
  • 4
  • Exam
  • 0,5
  • Total
  • 206,0
Continuous feedback during the course of the semester
Feedback by final exam (In addition to the grade)
Peer feedback (Students give each other feedback)

The students will be given oral feedback during the discussion confrontations and after the oral presentation. Written feedback will be given for the four exercise reports. These will also be followed up with oral, collective feedback. The students will also be given individual feedback to their exam essays and oral exam.

7,5 ECTS
Type of assessment
Written assignment, 5 page essay, 7 days
Oral examination, 20 minutes
The oral examination will consist of defence of the essay (10 minutes) and oral examination on the rest of the course contents (10 minutes). The examination is without preparation time. The final grade is given based on an overall assessment of the essay, the defence of the essay and the oral examination.
Exam registration requirements

Students must hand in at least three of the four exercise reports to take the exam.

Without aids
Marking scale
7-point grading scale
Censorship form
No external censorship
Two internal examiners

The same as ordinary exam. The written assignment must be reused at the re-exam. If the writtten assignment is not handed in as part of the ordinary exam the student must hand in an assignment based on a new topic provided by the teacher no later than three weeks before the re-exam.

If the requirement of handing in at least three of the four exercise reports is not fulfilled, the student must hand in the missing reports (based on data provided by the teachers) no later than three weeks before the reexam.  


Criteria for exam assesment

In order to obtain the grade 12 the student should convincingly and accurately demonstrate the knowledge, skills and competences described under Learning Outcome.